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Cellosaurus is database indexing available cell lines from various collections: American Type Culture Collection (ATCC), Bioresource Collection and Research Center, Taiwan (BCRC), Banco de Células do Rio de Janeiro (BCRJ), CellBank Australia (CBA), Collection of Cell Lines in Veterinary Medicine (CCLV), Cell Lines Service (CLS), Coriell, Drosophila Genomics Resource Center (DGRC), Deutsche Sammlung von Mikroorganismen und Zellkulturen GmbH (DSMZ), European Bank for induced pluripotent Stem Cells (EBiSC), European Collection of Authenticated Cell Cultures (ECACC), Interlab Cell Line Collection (ICLC), Institute for Fermentation Osaka (IFO), Istituto Zooprofilattico Sperimentale della Lombardia e dell'Emilia Romagna (IZSLER) biobank, Japanese Collection of Research Bioresources (JCRB) Cell Bank, Kunming Cell Bank of Type Culture Collection (KCB), Korean Cell Line Bank (KCLB), Millipore, National Cell Bank of Iran (NCBI_Iran), National Cancer Institute - Developmental Therapeutics Program (NCI-DTP), NINDS Human Cell and Data Repository (NHCDR), NIH AIDS Reagent Program (NIH-ARP), NISES, RIKEN Bioresource Center Cell Bank (RCB), Royan Stem Cell Bank (RSCB), Tick Cell Biobank (TCB), Tohoku University cell line catalog (TKG), and Ximbio.

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BW5147
 
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1+ mentions 		NCBI_Iran Cat# C542, RRID:CVCL_3896 Mus musculus (Mouse) Mouse lymphoma Group: Hybridoma fusion partner cell line. PMID:2530588
PMID:6970130
PMID:32419056 		Cancer cell line BW-5147, BW 5147 NCBI_Iran C542 BTO:BTO_0000791,
CLO:CLO_0002057,
CLDB:cl512,
Lonza:326,
NCBI_Iran:C542,
TKG:TKG 0729,
Wikidata:Q54798723		 CVCL_3896 2026-09-05 10:44:25 2
BV-173
 
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1+ mentions 		RRID:CVCL_0181 Homo sapiens (Human) Chronic myelogenous leukemia, BCR-ABL1 positive Population: Caucasian., Part of: LL-100 blood cancer cell line panel., Part of: COSMIC cell lines project., Part of: Cancer Dependency Map project (DepMap) (includes Cancer Cell Line Encyclopedia - CCLE). PMID:1720549
PMID:3098690
PMID:3332852
PMID:3856862
PMID:6572735
PMID:9680106
PMID:10071072
PMID:10576511
PMID:14504097
PMID:16523483
PMID:20164919
PMID:20215515
PMID:20809971
PMID:22460905
PMID:26589293
PMID:27397505
PMID:30285677
PMID:30629668
PMID:30894373
PMID:31068700
PMID:31160637
PMID:35839778 		Cancer cell line Male BV173 BTO:BTO_0004950,
CLO:CLO_0002052,
EFO:EFO_0002118,
MCCL:MCC:0000072,
CLDB:cl510,
CLDB:cl4968,
ArrayExpress:E-MTAB-38,
ArrayExpress:E-MTAB-783,
ArrayExpress:E-MTAB-2770,
ArrayExpress:E-MTAB-3610,
ArrayExpress:E-MTAB-7721,
ArrayExpress:E-MTAB-7722,
BioSample:SAMN03473373,
BioSample:SAMN10988488,
cancercelllines:CVCL_0181,
CCTCC:GDC0607,
Cell_Model_Passport:SIDM00962,
ChEMBL-Cells:CHEMBL3308228,
ChEMBL-Targets:CHEMBL2366145,
CLS:300133,
Cosmic:798672,
Cosmic:910710,
Cosmic:922164,
Cosmic:994184,
Cosmic:996292,
Cosmic:998702,
Cosmic:999741,
Cosmic:1019833,
Cosmic:1026570,
Cosmic:1037734,
Cosmic:1070701,
Cosmic:1071881,
Cosmic:1127249,
Cosmic:1176600,
Cosmic:1191703,
Cosmic:1524792,
Cosmic-CLP:910710,
DepMap:ACH-000432,
DSMZ:ACC-20,
DSMZCellDive:ACC-20,
EGA:EGAS00001000978,
GDSC:910710,
GEO:GSM236779,
GEO:GSM236815,
GEO:GSM886895,
GEO:GSM887960,
GEO:GSM1374414,
GEO:GSM1669633,
IARC_TP53:21199,
ICLC:HTL99018,
IGRhCellID:BV173,
LiGeA:CCLE_631,
LINCS_LDP:LCL-1111,
Lonza:117,
PharmacoDB:BV173_148_2019,
PRIDE:PXD030304,
Progenetix:CVCL_0181,
PubChem_Cell_line:CVCL_0181,
Wikidata:Q54798653		 CVCL_0181 2026-09-05 10:44:24 6
BVK168
 
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1+ mentions 		CCLV Cat# CCLV-RIE 1313, RRID:CVCL_A014 Myodes glareolus (Bank vole) Virology: Susceptible to infection by many bornaviruses, bunyaviruses, flaviviruses, poxviruses and togaviruses (PubMed=21729307). PMID:21729307
PMID:31513859 		Spontaneously immortalized cell line Male Bank Vole Kidney No. 168 CCLV CCLV-RIE 1313 CCLV:CCLV-RIE 1313,
Wikidata:Q54798681		 CVCL_A014 2026-09-05 10:44:25 1
BV-2
 
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1000+ mentions 		BCRJ Cat# 0356, RRID:CVCL_0182 Mus musculus (Mouse) PMID:2110186
PMID:19565166
PMID:21214938
PMID:21961028
PMID:22651808
PMID:26227174 		Transformed cell line Female BV2 BCRJ 0356 BTO:BTO_0003350,
EFO:EFO_0022792,
MCCL:MCC:0000073,
CLDB:cl7130,
BCRJ:0356,
BioGRID_ORCS_Cell_line:765,
CCRID:1101MOU-PUMC000063,
CCRID:4201MOU-CCTCC00311,
CCTCC:GDC0311,
ChEMBL-Cells:CHEMBL3307704,
ChEMBL-Targets:CHEMBL614781,
CLS:305156,
GEO:GSM635468,
GEO:GSM635469,
GEO:GSM635470,
GEO:GSM635471,
ICLC:ATL03001,
KCB:KCB 200770YJ,
Lonza:118,
PRIDE:PXD001597,
PRIDE:PXD004984,
PubChem_Cell_line:CVCL_0182,
Ubigene:YC-C035,
Wikidata:Q54798657		 CVCL_0182 2026-09-05 10:44:24 1029
BV-2
 
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1000+ mentions 		RRID:CVCL_0182 Mus musculus (Mouse) PMID:2110186
PMID:19565166
PMID:21214938
PMID:21961028
PMID:22651808
PMID:26227174 		Transformed cell line Female BV2 BTO:BTO_0003350,
EFO:EFO_0022792,
MCCL:MCC:0000073,
CLDB:cl7130,
BCRJ:0356,
BioGRID_ORCS_Cell_line:765,
CCRID:1101MOU-PUMC000063,
CCRID:4201MOU-CCTCC00311,
CCTCC:GDC0311,
ChEMBL-Cells:CHEMBL3307704,
ChEMBL-Targets:CHEMBL614781,
CLS:305156,
GEO:GSM635468,
GEO:GSM635469,
GEO:GSM635470,
GEO:GSM635471,
ICLC:ATL03001,
KCB:KCB 200770YJ,
Lonza:118,
PRIDE:PXD001597,
PRIDE:PXD004984,
PubChem_Cell_line:CVCL_0182,
Ubigene:YC-C035,
Wikidata:Q54798657		 CVCL_0182 2026-09-05 10:44:24 1029
BW5147
 
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1+ mentions 		TKG Cat# TKG 0729, RRID:CVCL_3896 Mus musculus (Mouse) Mouse lymphoma Group: Hybridoma fusion partner cell line. PMID:2530588
PMID:6970130
PMID:32419056 		Cancer cell line BW-5147, BW 5147 TKG TKG 0729 BTO:BTO_0000791,
CLO:CLO_0002057,
CLDB:cl512,
Lonza:326,
NCBI_Iran:C542,
TKG:TKG 0729,
Wikidata:Q54798723		 CVCL_3896 2026-09-05 10:44:25 2
BxPC3-R
 
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1+ mentions 		RRID:CVCL_XX78 Homo sapiens (Human) Pancreatic ductal adenocarcinoma Population: Caucasian. PMID:27229158
PMID:27910856 		Cancer cell line Female cancercelllines:CVCL_XX78,
GEO:GSM1906994,
GEO:GSM1906995,
GEO:GSM1906996,
Wikidata:Q93433506		 cvcl_0186 CVCL_XX78 2026-09-05 10:44:26 4
BXD-1425EPN
 
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1+ mentions 		RRID:CVCL_Y105 Homo sapiens (Human) Ependymoma Characteristics: Established from a second generation patient-derived xenograft established in the brain of RAG2/SCID mice. PMID:20511191 Cancer cell line Male BXD1425EPN, Baylor Xenograft Derived-1425EPN Wikidata:Q54798747 CVCL_Y105 2026-09-05 10:44:26 1
BY00867
 
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1+ mentions 		RRID:CVCL_9T50 Homo sapiens (Human) Part of: International Histocompatibility Workshop cell lines. PMID:22077627 Transformed cell line Sex unspecified IPD-IMGT/HLA:25491,
Wikidata:Q54799663		 CVCL_9T50 2026-09-05 10:44:34 1
C-20/A4
 
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1+ mentions 		Millipore Cat# SCC041, RRID:CVCL_6821 Homo sapiens (Human) PMID:7989586
PMID:15604530 		Transformed cell line Male C-20\A4, C20/A4, C-20a4, C20A4 Millipore SCC041 BTO:BTO_0005670,
ChEMBL-Cells:CHEMBL3308083,
ChEMBL-Targets:CHEMBL613871,
Millipore:SCC041,
PubChem_Cell_line:CVCL_6821,
Wikidata:Q54799734		 CVCL_6821 2026-09-05 10:44:34 1
c-WRT-7-LR
 
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1+ mentions
Discontinued 		RRID:CVCL_J367 Breed/subspecies: Wistar King Aptekman/Hok (WKA/Hok)., Transformant: NCBI_TaxID; 11938; Rauscher murine leukemia virus (R-MuLV)., Discontinued: JCRB; FDSC0004; true. PMID:3469020 Cancer cell line Female LR JCRB:JCRB0168,
JCRB:FDSC0004,
Wikidata:Q54799771		 CVCL_J367 2026-09-05 10:44:35 2
C-20/A4
 
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1+ mentions 		RRID:CVCL_6821 Homo sapiens (Human) PMID:7989586
PMID:15604530 		Transformed cell line Male C-20\A4, C20/A4, C-20a4, C20A4 BTO:BTO_0005670,
ChEMBL-Cells:CHEMBL3308083,
ChEMBL-Targets:CHEMBL613871,
Millipore:SCC041,
PubChem_Cell_line:CVCL_6821,
Wikidata:Q54799734		 CVCL_6821 2026-09-05 10:44:34 1
C-26
 
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1+ mentions 		RRID:CVCL_XC68 Mus musculus (Mouse) Group: Patented cell line. Hybridoma C26 Wikidata:Q93436332 CVCL_XC68 2026-09-05 10:44:34 2
C-4-I
 
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1+ mentions
Discontinued 		ATCC Cat# CRL-7396, RRID:CVCL_2253 Homo sapiens (Human) Human papillomavirus-related cervical squamous cell carcinoma Virology: Contains a complete HPV18 genome., Population: Caucasian., Part of: Naval Biosciences Laboratory (NBL) collection (transferred to ATCC in 1982)., Part of: COSMIC cell lines project., Part of: Cancer Dependency Map project (DepMap) (includes Cancer Cell Line Encyclopedia - CCLE). PMID:77569
PMID:327080
PMID:560251
PMID:833871
PMID:2983228
PMID:2990217
PMID:3029760
PMID:3518877
PMID:5796380
PMID:6935474
PMID:7459858
PMID:12661003
PMID:13863218
PMID:14114965
PMID:15531914
PMID:20215515
PMID:27397505
PMID:29156801
PMID:30894373
PMID:35692560
PMID:35839778 		Cancer cell line Female C-4I, C-4 I, C-4-i, C4-I, C4-1, C4 I, C4I, C41, Hs 636.T, Hs 636 T ATCC CRL-7396 BTO:BTO_0006016,
CLO:CLO_0002125,
CLO:CLO_0002129,
EFO:EFO_0002122,
CLDB:cl521,
ArrayExpress:E-MTAB-38,
ArrayExpress:E-MTAB-3610,
ATCC:CRL-1594,
ATCC:CRL-7396,
BioSample:SAMN03471401,
cancercelllines:CVCL_2253,
Cell_Model_Passport:SIDM00905,
Cosmic:687506,
Cosmic:877471,
Cosmic:925358,
Cosmic:928906,
Cosmic:1019308,
Cosmic:1131518,
Cosmic:1193035,
Cosmic:1201771,
Cosmic:2646656,
Cosmic:2660189,
Cosmic:2816213,
Cosmic-CLP:687506,
DepMap:ACH-001334,
ECACC:89051702,
EGA:EGAS00001000978,
GDSC:687506,
GEO:GSM113860,
GEO:GSM253295,
GEO:GSM675731,
GEO:GSM685438,
GEO:GSM1669639,
IGRhCellID:C4I,
LINCS_LDP:LCL-1301,
PharmacoDB:C4I_151_2019,
PRIDE:PXD030304,
Progenetix:CVCL_2253,
Wikidata:Q54799744		 CVCL_2253 2026-09-05 10:44:35 1
C-4-II
 
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1+ mentions
Discontinued 		ATCC Cat# CRL-1595, RRID:CVCL_1095 Homo sapiens (Human) Human papillomavirus-related cervical squamous cell carcinoma Virology: Contains a complete HPV18 genome., Population: Caucasian., Part of: Naval Biosciences Laboratory (NBL) collection (transferred to ATCC in 1982)., Part of: Cancer Dependency Map project (DepMap) (includes Cancer Cell Line Encyclopedia - CCLE). PMID:833871
PMID:2990217
PMID:3518877
PMID:5796380
PMID:6935474
PMID:13863218
PMID:14114965
PMID:15531914
PMID:20164919
PMID:20215515
PMID:24134916
PMID:29156801
PMID:35692560 		Cancer cell line Female C-4 II, C-4II, C4 II, C4II ATCC CRL-1595 CLO:CLO_0002126,
EFO:EFO_0002123,
CLDB:cl571,
ArrayExpress:E-MTAB-38,
ArrayExpress:E-MTAB-783,
ATCC:CRL-1595,
ATCC:CRL-7914,
BioSample:SAMN03471403,
cancercelllines:CVCL_1095,
Cell_Model_Passport:SIDM00906,
ChEMBL-Cells:CHEMBL3308712,
ChEMBL-Targets:CHEMBL1075400,
Cosmic:760218,
Cosmic:809230,
Cosmic:910553,
DepMap:ACH-001335,
ECACC:88102701,
GEO:GSM675732,
GEO:GSM685437,
IARC_TP53:21202,
IGRhCellID:C4II,
LINCS_LDP:LCL-1302,
Progenetix:CVCL_1095,
PubChem_Cell_line:CVCL_1095,
Wikidata:Q54799745		 CVCL_1095 2026-09-05 10:44:35 3
C-4-II
 
Resource Report
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1+ mentions
Discontinued 		RRID:CVCL_1095 Homo sapiens (Human) Human papillomavirus-related cervical squamous cell carcinoma Virology: Contains a complete HPV18 genome., Population: Caucasian., Part of: Naval Biosciences Laboratory (NBL) collection (transferred to ATCC in 1982)., Part of: Cancer Dependency Map project (DepMap) (includes Cancer Cell Line Encyclopedia - CCLE). PMID:833871
PMID:2990217
PMID:3518877
PMID:5796380
PMID:6935474
PMID:13863218
PMID:14114965
PMID:15531914
PMID:20164919
PMID:20215515
PMID:24134916
PMID:29156801
PMID:35692560 		Cancer cell line Female C-4 II, C-4II, C4 II, C4II CLO:CLO_0002126,
EFO:EFO_0002123,
CLDB:cl571,
ArrayExpress:E-MTAB-38,
ArrayExpress:E-MTAB-783,
ATCC:CRL-1595,
ATCC:CRL-7914,
BioSample:SAMN03471403,
cancercelllines:CVCL_1095,
Cell_Model_Passport:SIDM00906,
ChEMBL-Cells:CHEMBL3308712,
ChEMBL-Targets:CHEMBL1075400,
Cosmic:760218,
Cosmic:809230,
Cosmic:910553,
DepMap:ACH-001335,
ECACC:88102701,
GEO:GSM675732,
GEO:GSM685437,
IARC_TP53:21202,
IGRhCellID:C4II,
LINCS_LDP:LCL-1302,
Progenetix:CVCL_1095,
PubChem_Cell_line:CVCL_1095,
Wikidata:Q54799745		 CVCL_1095 2026-09-05 10:44:35 3
C-1300 clone N18
 
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1+ mentions 		TKG Cat# TKG 0307, RRID:CVCL_4724 Mus musculus (Mouse) Mouse neuroblastoma PMID:2759161
PMID:4400294
PMID:5283932
PMID:21214938 		Cancer cell line Male N-18, N18, N18 (Hamprecht), C-1300N18 TKG TKG 0307 CLO:CLO_0007920,
CLO:CLO_0050185,
CLDB:cl3614,
ECACC:08062527,
GEO:GSM635472,
GEO:GSM635473,
GEO:GSM635474,
GEO:GSM635475,
RCB:RCB2620,
TKG:TKG 0307,
Wikidata:Q54907320		 cvcl_4343 CVCL_4724 2026-09-05 10:44:34 1
C0135
 
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1+ mentions 		RRID:CVCL_K404 Homo sapiens (Human) Population: Caucasian., Part of: ECACC randomly selected UK Caucasian blood donors cell line selection. Transformed cell line Female ECACC:93021107,
Wikidata:Q54799955		 CVCL_K404 2026-09-05 10:44:39 2
CaR-1
 
Resource Report
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1+ mentions 		RRID:CVCL_1116 Homo sapiens (Human) Rectal adenocarcinoma Caution: TP53 mutation indicated incorrectly as being at c.701A>G in PubMed=9290701., Population: Japanese., Part of: COSMIC cell lines project., Part of: Cancer Dependency Map project (DepMap) (includes Cancer Cell Line Encyclopedia - CCLE)., Part of: AstraZeneca Colorectal cell line (AZCL) panel. PMID:9290701
PMID:16418264
PMID:18258742
PMID:20164919
PMID:20215515
PMID:20606684
PMID:23272949
PMID:25485619
PMID:25877200
PMID:25926053
PMID:26589293
PMID:27397505
PMID:28854368
PMID:29101300
PMID:30894373
PMID:35839778 		Cancer cell line Male CaR1, CAR1 BTO:BTO_0003131,
CLO:CLO_0009955,
EFO:EFO_0006548,
ArrayExpress:E-MTAB-783,
ArrayExpress:E-MTAB-2706,
ArrayExpress:E-MTAB-3610,
BioSample:SAMN03472863,
cancercelllines:CVCL_1116,
Cell_Model_Passport:SIDM00488,
ChEMBL-Cells:CHEMBL3308100,
ChEMBL-Targets:CHEMBL1075418,
ColonAtlas:CAR1,
Cosmic:873697,
Cosmic:924108,
Cosmic:1122321,
Cosmic:1184086,
Cosmic:1187300,
Cosmic-CLP:924108,
DepMap:ACH-002345,
EGA:EGAS00001000610,
EGA:EGAS00001000978,
GDSC:924108,
GEO:GSM827225,
GEO:GSM1374441,
GEO:GSM1448170,
GEO:GSM1669665,
IARC_TP53:21665,
IARC_TP53:21735,
JCRB:JCRB0207,
LINCS_LDP:LCL-1288,
PharmacoDB:CaR1_186_2019,
PRIDE:PXD005235,
PRIDE:PXD005354,
PRIDE:PXD005355,
PRIDE:PXD030304,
Progenetix:CVCL_1116,
PubChem_Cell_line:CVCL_1116,
Wikidata:Q54808598		 CVCL_1116 2026-09-05 10:45:18 4
Capan-2
 
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100+ mentions 		RRID:CVCL_0026 Homo sapiens (Human) Pancreatic ductal adenocarcinoma Caution: TP53 mutation indicated incorrectly as being at c.818G>A in PubMed=1630814., Population: Caucasian., From: Memorial Sloan Kettering Cancer Center; New York; USA., Part of: NCI RAS program mutant KRAS cell line panel., Part of: Human Protein Atlas, Cell Atlas panel., Part of: COSMIC cell lines project., Part of: Cancer Dependency Map project (DepMap) (includes Cancer Cell Line Encyclopedia - CCLE). PMID:1630814
PMID:1764370
PMID:3019537
PMID:3518877
PMID:6582512
PMID:6935474
PMID:7459858
PMID:7809022
PMID:7961102
PMID:7972006
PMID:8026879
PMID:8194712
PMID:8286197
PMID:8426738
PMID:10027410
PMID:11169957
PMID:11169959
PMID:11668190
PMID:12692724
PMID:12800145
PMID:14695172
PMID:15126341
PMID:15367885
PMID:15688027
PMID:15770730
PMID:16957166
PMID:18298655
PMID:18380791
PMID:20037478
PMID:20164919
PMID:20215515
PMID:20418756
PMID:22460905
PMID:22585861
PMID:24700732
PMID:25167228
PMID:25485619
PMID:25877200
PMID:26216984
PMID:26589293
PMID:27259358
PMID:27397505
PMID:30894373
PMID:31037374
PMID:31068700
PMID:35839778 		Cancer cell line Male CaPan-2, CAPAN-2, Capan 2, CAPAN 2, Capan2, CAPAN2 BTO:BTO_0002745,
CLO:CLO_0002198,
EFO:EFO_0002154,
MCCL:MCC:0000127,
CLDB:cl642,
CLDB:cl643,
CLDB:cl644,
ArrayExpress:E-MTAB-38,
ArrayExpress:E-MTAB-783,
ArrayExpress:E-MTAB-2706,
ArrayExpress:E-MTAB-2770,
ArrayExpress:E-MTAB-3610,
ATCC:HTB-80,
BCRJ:0060,
BioSample:SAMN03472025,
BioSample:SAMN03473387,
BioSample:SAMN10988212,
cancercelllines:CVCL_0026,
CancerTools:161908,
Cell_Model_Passport:SIDM00943,
CGH-DB:163-1,
CGH-DB:9270-4,
ChEMBL-Cells:CHEMBL3308720,
ChEMBL-Targets:CHEMBL1075417,
CLS:300144,
Cosmic:730527,
Cosmic:736271,
Cosmic:755303,
Cosmic:808163,
Cosmic:868246,
Cosmic:910915,
Cosmic:913303,
Cosmic:922256,
Cosmic:923167,
Cosmic:932517,
Cosmic:932987,
Cosmic:933522,
Cosmic:934554,
Cosmic:947396,
Cosmic:948727,
Cosmic:949232,
Cosmic:1006366,
Cosmic:1108339,
Cosmic:1299297,
Cosmic:1320463,
Cosmic:1366277,
Cosmic:1571775,
Cosmic:1995363,
Cosmic:2009527,
Cosmic:2046535,
Cosmic:2434087,
Cosmic:2668288,
Cosmic-CLP:910915,
DepMap:ACH-000107,
DSMZ:ACC-245,
DSMZCellDive:ACC-245,
EGA:EGAS00001000610,
EGA:EGAS00001000978,
GDSC:910915,
GEO:GSM206453,
GEO:GSM621896,
GEO:GSM784710,
GEO:GSM886921,
GEO:GSM887987,
GEO:GSM1024399,
GEO:GSM1374440,
GEO:GSM1669664,
GEO:GSM7701427,
GEO:GSM7701428,
IARC_TP53:313,
IARC_TP53:21042,
IARC_TP53:30199,
IGRhCellID:Capan2,
IZSLER:BS TCL 10,
KCLB:30080,
LiGeA:CCLE_178,
LINCS_LDP:LCL-1733,
Lonza:1464,
MetaboLights:MTBLS812,
PharmacoDB:Capan2_185_2019,
PRIDE:PXD003198,
PRIDE:PXD030304,
Progenetix:CVCL_0026,
PubChem_Cell_line:CVCL_0026,
SKY/M-FISH/CGH:1993,
Wikidata:Q54808591		 CVCL_0026 2026-09-05 10:45:17 298

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The NIDDK Information Network (dkNET) serves the needs of basic and clinical investigators by providing seamless access to large pools of data and research resources relevant to the mission of The National Institute of Diabetes Digestive and Kidney Diseases (NIDDK). dkNET is hosted at University of California San Diego, and is supported by NIH NIDDK grant U24DK097771.

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